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Enjoying their normal lives, Jeremy Belpois, Ulrich Stern, William Dunbar, Odd Della Robbia, Yumi Ishiyama and Aelita Shaffer soon discover that there is trouble still brewing at Kadic Academy. Even worse, they discover that their old and greatest virtual enemy, the evil, sentient artificial intelligence X.A.N.A. is somehow back and even stronger and more dangerous than ever before, as a highly evolved and modified version of itself.
X.A.N.A now wants to take back his energy-source codes that were implanted in the minds of Aelita, Odd, Ulrich, and Yumi during the time of his supposed "destruction" in order to gain control over the world network once again. The war against X.A.N.A is far from over, and the five Lyoko warriors are back in action.